Mammals (formally Mammalia) are a class The composition of each class is determined by a taxonomist. Often there is no exact agreement, with different taxonomists taking different positions. There are no hard rules that a taxonomist needs to follow in describing a class, but for well-known animals there is likely to be consensus. For example, dogs are usually assigned to the phylum of vertebrate Vertebrates are members of the subphylum Vertebrata, chordates with backbones and spinal columns. About 58,000 species of vertebrates have been currently described. Vertebrata is the largest subphylum of chordates, and contains many familiar groups of large land animals. Vertebrates are comprised of animals from the groups jawless fishs, bony fish,, air-breathing animals Animals are a major group of mostly multicellular, eukaryotic organisms of the kingdom Animalia or Metazoa. Their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later on in their life. Most animals are motile, meaning they can move spontaneously and independently. All animals are also whose females are characterized by the possession of mammary glands Mammary glands are the organs that, in mammals, produce milk for the sustenance of the young. These exocrine glands are enlarged and modified sweat glands and give mammals their name. The mammary glands of domestic mammals that have more than two breasts are called dugs while both males and females are characterized by hair Hair is a filamentous biomaterial, that grows from follicles found in the dermis. The human body, apart from its glabrous skin, is covered in follicles which produce thick terminal and fine vellus hair. Most common interest in hair is focused on hair growth, hair types and hair care, but hair is also an important biomaterial primarily composed of and/or fur, three middle ear The middle ear is the portion of the ear internal to the eardrum, and external to the oval window of the cochlea. The mammalian middle ear contains three ossicles, which couple vibration of the eardrum into waves in the fluid and membranes of the inner ear. The hollow space of the middle ear has also been called the tympanic cavity, or cavum bones Bones are rigid organs that form part of the endoskeleton of vertebrates. They function to move, support, and protect the various organs of the body, produce red and white blood cells and store minerals. Bone tissue is a type of dense connective tissue. Because bones come in a variety of shapes and have a complex internal and external structure used in hearing Hearing is one of the traditional five senses. It is the ability to perceive sound by detecting vibrations via an organ such as the ear. The inability to hear is called deafness, and a neocortex The neocortex , also called the neopallium ("new mantel") and isocortex ("equal rind"), is a part of the brain of mammals. It is the outer layer of the cerebral hemispheres, and made up of six layers, labelled I to VI (with VI being the innermost and I being the outermost). The neocortex is part of the cerebral cortex (along region in the brain. Some mammals have sweat glands 1. Hair Shaft 2. Stratum Corneum 3. Pigment Layer 4. Stratum Spinosum 5. Stratum Basale 6. Arrector Pili Muscle 7. Sebaceous Gland 8. Hair Follicle 9. Papilla of Hair 10. Nerve Fiber 11. Sweat Gland 12. Pacinian Corpuscle 13. Artery 14. Vein 15. Sensory Nerve ending 16. Dermal Papillary 17. Sweat Pore, but most do not.

Mammals are divided into three main infraclass taxa depending how they are born. These taxa are: monotremes Monotremes are mammals that lay eggs (Prototheria) instead of giving birth to live young like marsupials (Metatheria) and placental mammals (Eutheria), marsupials Marsupials are an infraclass of mammals, characterized by a distinctive pouch , in which females carry their young through early infancy. In modern times, they are well-known for being the dominant group of mammals in Australia, though there are also a number of species found in the Americas, as well as on the island of New Guinea and placentals Eutheria are a group of mammals consisting of placental mammals plus all extinct mammals that are more closely related to living placentals (such as humans) than to living marsupials (such as kangaroos). They are distinguished from non-eutherians by various features of the feet, ankles, jaws and teeth. One of the major differences between. Except for the five species of monotremes Monotremes are mammals that lay eggs (Prototheria) instead of giving birth to live young like marsupials (Metatheria) and placental mammals (Eutheria) (which lay eggs), all mammal species give birth to live young. Most mammals also possess specialized teeth Teeth are small, calcified, whitish structures found in the jaws (or mouths) of many vertebrates that are used to break down food. Some animals, particularly carnivores, also use teeth for hunting or for defensive purposes. The roots of teeth are covered by gums. Teeth are not made of bone, but rather of multiple tissues of varying density and, and the largest group of mammals, the placentals Eutheria are a group of mammals consisting of placental mammals plus all extinct mammals that are more closely related to living placentals (such as humans) than to living marsupials (such as kangaroos). They are distinguished from non-eutherians by various features of the feet, ankles, jaws and teeth. One of the major differences between, use a placenta The placenta is an organ that connects the developing fetus to the uterine wall to allow nutrient uptake, waste elimination, and gas exchange via the mother's blood supply. Placentas are a defining characteristic of eutherian or "placental" mammals, but are also found in some snakes and lizards with varying levels of development up to during gestation. The mammalian brain regulates endothermic In biology, a warm-blooded animal species is one whose members maintain thermal homeostasis; that is, they keep their body temperature at a roughly constant level, regardless of the ambient temperature. This involves the ability to cool down or produce more body heat. Warm-blooded animals mainly control their body temperature by regulating their and circulatory The circulatory system is an organ system that passes nutrients , gases, hormones, blood cells, etc. to and from cells in the body to help fight diseases and help stabilize body temperature and pH to maintain homeostasis systems, including a four-chambered heart The heart is a myogenic muscular organ found in all animals with a circulatory system , that is responsible for pumping blood throughout the blood vessels by repeated, rhythmic contractions. The term cardiac (as in cardiology) means "related to the heart" and comes from the Greek καρδιά, kardia, for "heart".

There are approximately 5,400 species In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are often used, such as similarity of DNA, morphology or of mammals, distributed in about 1,200 genera In biology, a genus is a low-level taxonomic rank (a taxon) used in the classification of living and fossil organisms, which is an example of definition by genus and differentia. The term comes from Latin genus "descent, family, type, gender", cognate with Greek: γένος – genos, "race, stock, kin", 153 families What does and does not belong to each family is determined by a taxonomist. Similarly for the question if a particular family should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing a family, and 29 orders The Latin suffix -formes meaning "having the form of" is used for the scientific name of orders of birds and fishes, but not for those of mammals and invertebrates[1] (though this varies by classification scheme Biological classification, or scientific classification in biology, is a method by which biologists group and categorize organisms by biological type, such as genus or species. Biological classification is a form of scientific taxonomy, but should be distinguished from folk taxonomy, which lacks scientific basis. Modern biological classification). Mammals range in size from the 30–40 millimeter (1- to 1.5-inch) Bumblebee Bat Kitti's Hog-nosed Bat , also known as the bumblebee bat, is a vulnerable species of bat and the only member of the family Craseonycteridae. It occurs in western Thailand and southeast Burma, where it occupies limestone caves along rivers to the 33-meter (108-foot) Blue Whale The Blue whale is a marine mammal belonging to the suborder of baleen whales (called Mysticeti). At over 33 metres (108 ft) in length and 180 metric tons (200 short tons) or more in weight, it is the largest animal ever known to have existed.

Mammals are divided into two subclasses: the Prototheria Prototheria is a taxonomic group, or taxon, to which the order Monotremata belongs. It is conventionally ranked as a subclass within the mammals, which includes the oviparous monotremes, and the Theria Theria is a subclass of mammals that give birth to live young without using a shelled egg, including both eutherians (placental mammals) and metatherians (marsupials and their ancestors).[clarification needed], which includes the placentals and live-bearing marsupials. Most mammals, including the six largest orders, belong to the placental Eutheria are a group of mammals consisting of placental mammals plus all extinct mammals that are more closely related to living placentals (such as humans) than to living marsupials (such as kangaroos). They are distinguished from non-eutherians by various features of the feet, ankles, jaws and teeth. One of the major differences between group. The three largest orders, in descending order, are Rodentia Rodentia is an order of mammals also known as rodents, characterised by two continuously growing incisors in the upper and lower jaws which must be kept short by gnawing (mice, rats, porcupines, beavers, capybaras, and other gnawing mammals), Chiroptera Bats are flying mammals in the order Chiroptera . The forelimbs of bats are webbed and developed as wings, making them the only mammals naturally capable of true and sustained flight. By contrast, other mammals said to fly, such as flying squirrels, gliding possums and colugos, glide rather than fly, and can only glide for short distances. Bats do (bats), and Soricomorpha The order Soricomorpha is a biological clade within the class of mammals. In previous years it formed a significant group within the former Insectivora order. However, that order was shown to be polyphyletic and various new orders were split off from it, including Afrosoricida (tenrecs and golden moles), Macroscelidea (elephant shrews), and (shrews, moles and solenodons). The next three largest orders include the Carnivora The diverse order Carnivora (pronounced /kɑrˈnɪvərə/ or sometimes /ˌkɑrnɪˈvɔərə/; from Latin carō "flesh", + vorāre "to devour") includes over 260 species of placental mammals. Its members are formally referred to as carnivorans, while the word "carnivore" (often popularly applied to members of this (dogs The dog is a domesticated form of the gray wolf, a member of the Canidae family of the order Carnivora. The term is used for both feral and pet varieties. The domestic dog has been one of the most widely kept working, hunting and companion animals in human history, cats The cat , also known as the domestic cat or housecat to distinguish it from other felines and felids, is a small domesticated carnivorous mammal that is valued by humans for its companionship and ability to hunt vermin and household pests. Cats have been associated with humans for at least 9,500 years, and are currently the most popular pet in the, weasels, bears, seals, and their relatives), the Cetartiodactyla Cetartiodactyla is the clade to which whales and even-toed ungulates have currently been placed. The term was coined by merging the name for the two orders, Cetacea and Artiodactyla, into a single word. The term Cetartiodactyla reflects the idea that whales evolved from within the artiodactyls. Under this definition their closest living land (including the even-toed hoofed mammals and the whales) and the Primates A primate is a member of the biological order Primates (/praɪˈmeɪtiːz/ prī·mā′·tēz; Latin: "prime, first rank"), the group that contains prosimians (including lemurs, lorises, galagos and tarsiers ) and simians (monkeys and apes). With the exception of humans, who inhabit every continent on Earth,[a] most primates live in to which the human Humans, known taxonomically as Homo sapiens , are the only living species in the Homo genus of bipedal primates in Hominidae, the great ape family. However, in some cases "human" is used to refer to any member of the genus Homo species belongs. The relative size of these latter three orders differs according to the classification scheme and definitions used by various authors.

Phylogenetically In biology, phylogenetics is the study of evolutionary relatedness among various groups of organisms , which is discovered through molecular sequencing data and morphological data matrices. The term phylogenetics is of Greek origin from the terms phyle/phylon (φυλή/φῦλον), meaning "tribe, race," and genetikos (γενετικός, Mammalia is defined as all descendants of the most recent common ancestor In genetics, the most recent common ancestor of any set of organisms is the most recent individual from which all organisms in the group are directly descended. The term is often applied to human genealogy of monotremes (e.g., echidnas Echidnas , also known as spiny anteaters, belong to the family Tachyglossidae in the monotreme order of egg-laying mammals. There are four extant species, which, together with the Platypus, are the only surviving members of that order and are the only extant mammals that lay eggs. Although their diet consists largely of ants and termites, they are and platypuses The Platypus is a semi-aquatic mammal endemic to eastern Australia, including Tasmania. Together with the four species of echidna, it is one of the five extant species of monotremes, the only mammals that lay eggs instead of giving birth to live young. It is the sole living representative of its family (Ornithorhynchidae) and genus () and therian mammals (marsupials Marsupials are an infraclass of mammals, characterized by a distinctive pouch , in which females carry their young through early infancy. In modern times, they are well-known for being the dominant group of mammals in Australia, though there are also a number of species found in the Americas, as well as on the island of New Guinea and placentals).[2] This means that some extinct groups of "mammals" are not members of the crown group A crown group is the smallest monophyletic group, or "clade", to contain the last common ancestor of all extant members, and all of that ancestor's descendants. Extinct organisms can still be part of a crown group: for instance, the extinct dodo is still descended from the last common ancestor of all living birds, so falls within the Mammalia, even though most of them have all the characteristics that traditionally would have classified them as mammals.[3] These "mammals" are now usually placed in the unranked clade Mammaliaformes Mammaliaformes is a clade that contains the mammals and their closest extinct relatives. Phylogenetically it is defined as a clade including the most recent common ancestor of Sinoconodon, morganuconodonts, docodonts, Monotremata, Marsupialia, Placentalia, extinct members of this clade, and all of its descendants. The precise phylogeny is disputed.

The mammalian line of descent diverged from an amniote line at the end of the Carboniferous The Carboniferous is a geologic period and system that extends from the end of the Devonian Period, about 359.2 ± 2.5 Ma , to the beginning of the Permian Period, about 299.0 ± 0.8 Ma (ICS, 2004, period. One line of amniotes would lead to reptiles Reptiles are animals in the class Reptilia. They are characterized by breathing air, laying tough-shelled amniotic eggs, and having skin covered in scales and/or scutes. Reptiles are classically viewed as having a "cold-blooded" metabolism. They are tetrapods (either having four limbs or being descended from four-limbed ancestors), while the other would lead to synapsids Synapsids are a group of animals that includes mammals and everything more closely related to mammals than to other living amniotes. Primitive synapsids are usually called pelycosaurs; more advanced mammal-like ones, therapsids. The non-mammalian members are described as mammal-like reptiles in classical systematics, but are referred to as ". According to cladistics Cladistics is a method of classifying species of organisms into groups called clades, which consist of 1) all the descendants of an ancestral organism and 2) the ancestor itself. For example, birds, dinosaurs, crocodiles, and all descendants of their most recent common ancestor form a clade. In the terms of biological systematics, a clade is a, mammals are a sub-group of synapsids. Although they were preceded by many diverse groups of non-mammalian synapsids (sometimes misleadingly referred to as mammal-like reptiles), the first true mammals appeared in the Triassic period. Modern mammalian orders appeared in the Palaeocene and Eocene epochs of the Palaeogene period.

Contents

Distinguishing features

Living mammal species can be identified by the presence of sweat glands, including those that are specialized to produce milk. However, other features are required when classifying fossils, since soft tissue glands and some other features are not visible in fossils. Paleontologists use a distinguishing feature that is shared by all living mammals (including monotremes), but is not present in any of the early Triassic synapsids: mammals use two bones for hearing that were used for eating by their ancestors. The earliest synapsids had a jaw joint composed of the articular (a small bone at the back of the lower jaw) and the quadrate (a small bone at the back of the upper jaw). Most reptiles including lizards, crocodilians, dinosaurs (and their descendants the birds) use this system, as did non-mammalian synapsids such as therapsids. Mammals have a different jaw joint, however, composed only of the dentary (the lower jaw bone which carries the teeth) and the squamosal (another small skull bone). In mammals the quadrate and articular bones have become the incus and malleus bones in the middle ear.

Mammals also have a double occipital condyle: they have two knobs at the base of the skull which fit into the topmost neck vertebra, and other vertebrates have a single occipital condyle. Paleontologists use only the jaw joint and middle ear as criteria for identifying fossil mammals, since it would be confusing if they found a fossil that had one feature, but not the other.

Classification

Main article: Mammal classification Over 70% of mammal species are in the orders Rodentia (blue), Chiroptera (red), and Soricomorpha (yellow)

George Gaylord Simpson's "Principles of Classification and a Classification of Mammals" (AMNH Bulletin v. 85, 1945) was the original source for the taxonomy listed here. Simpson laid out a systematics of mammal origins and relationships that was universally taught until the end of the 20th century. Since Simpson's classification, the paleontological record has been recalibrated, and the intervening years have seen much debate and progress concerning the theoretical underpinnings of systematization itself, partly through the new concept of cladistics. Though field work gradually made Simpson's classification outdated, it remained the closest thing to an official classification of mammals.

Standardized textbook classification

A somewhat standardized classification system has been adopted by most current mammalogy classroom textbooks. The following taxonomy of extant and recently extinct mammals is from Vaughan et al. (2000).

Class Mammalia

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